Original silver on the wheels looks much better. Same thing for the GT2 on the pic.
IMO, painted wheels very rarely work and when they do it's only for a specific design and usually painted black or dark grey (on black or silver cars) -- even then for not much extra effect. One more comment, amber lights actually look quite nice (and warm) with the (warm) brown interior and the green body on your Box.
But taste is personal. If it looks better to you, paint them.
Z.
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