Great progress!
When you connected the RCA line to the speaker, you were connecting a 'pre-amp' (Low voltage tiny power) wire to the speaker. The RCA connection does not have the capacity to drive a speaker directly...it would have to go to an amp and then the amp connects to the crossover. The output from your new radio was 'line level', or amplified, hence your success driving the speakers.
The different connections for the tweeter crossover output will determine the 'brightness' of the treble. Connect to whichever one sounds best to you.
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